The Nittany Lions allowed 260 yards, one field goal, and a garbage-time touchdown in the 27-11 victory.
The Nittany Lions managed just 85 rushing yards without their leading rusher.
Warren broke the program record for most touchdowns by a tight end, while Clifford had his first career 100-yard game.
Singleton was noticeably absent from practice on Wednesday.
Here's everything you need to know before the Stripe Out this Saturday.
The Bruins have only beaten Hawaii this season.