Collins did not play in her first year with the Nittany Lions.
Singleton leaves Happy Valley as the all-time leader in total and rushing touchdowns.
Penn State football continues to make changes to their staff.
Matt Campbell continues to fill out his new staff.
Mouser spent 10 seasons with Campbell in Ames, Iowa, and spent the past two seasons as the Cyclones' play caller.
Before Clanton joined Campbell's staff in Ames, the Cyclones averaged just 108 yards per game.