The Nittany Lions gave up 11 unanswered goals in the first half.
Knowles was announced as Penn State's next defensive coordinator on January 27.
What the two Nittany Lions now have in common: Penn State degrees, Super Bowl rings, and Matty Fresh songs.
The Joe Humphreys Fly Fishing program has a storied history at Penn State.
The Nittany Lions have dropped seven of their last eight games and are in danger of missing the Big Ten Tournament.
Penn State Athletics reported $5.6 million in profit for the 2024 fiscal year.
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