About 2.6% of tests administered this week returned positive results.
Bolton leaves an Iowa State squad that went 2-22 this past season.
The freshly drafted attackman is the first Nittany Lion to be given three All-Big Ten first-team honors in program history
Penn State men's basketball added another piece to the 2021-22 schedule on Thursday.
Oweh leaves Penn State with 63 tackles and seven sacks over three seasons.
After a whirlwind of first-round trades, Parsons ended up in his dream situation in Dallas.