Penn State had its six-game win streak snapped after turning the ball over 17 times.
Penn State remains atop its conference after its weekend sweep.
Penn State fell just short of a comeback win in its season opener at Panzer Stadium.
This was the first win in Bloomington for the Nittany Lions since 2014.
The Nittany Lions hit .433 along with racking up six aces and double-digit kills from Michael Valenzi, Michal Kowal, and John Kerr.
Penn State female fall and spring sports will be celebrated over the next week to commemorate NGWSD, which is officially held on Wednesday, February 7.