Lane played every game for the Nittany Lions over the past two seasons.
Boyer notched his first career start in the Pinstripe Bowl.
The redshirt freshman appeared in four games for the Nittany Lions.
The freshman appeared in two games for Penn State this season.
The true freshman had 16 tackles in 10 games.
Ikakoula adds to a growing list of Iowa State transfers following Matt Campbell's hiring.