Eskildsen had 30 receptions for 526 yards and five touchdowns in 2025.
Manske is the sixth former Iowa State player to transfer to Penn State, following fellow quarterback Rocco Becht.
Becht leaves Iowa State with 9,274 passing yards.
Burkle has one year of eligibility remaining.
Artis didn't see any action in 2025.
Neal is the third transfer to commit to Penn State.
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