Warren had another record-breaking day against the Trojans.
Warren now holds the single-game school record for most receptions by an offensive player, passing DaeSean Hamilton who had 14 receptions in a game in 2014.
The Nittany Lions mounted a comeback bid in the third period but fell short.
Ryan Barker's walk-off overtime field goal completed Penn State's comeback.
The Nittany Lions secured their first series sweep of the season against Union.