Twenty-one attack errors and 23 service errors was the Achilles' heel of the Nittany Lions.
Penn State had its six-game win streak snapped after turning the ball over 17 times.
Penn State remains atop its conference after its weekend sweep.
Penn State fell just short of a comeback win in its season opener at Panzer Stadium.
This was the first win in Bloomington for the Nittany Lions since 2014.
Johnny Dixon, Adisa Isaac, Kalen King, and Theo Johnson will represent the Nittany Lions.