Reed tallied 181 total tackles, 12.5 tackles for loss, 3.5 sacks, 12 pass deflections, and five interceptions in 50 appearances for the Nittany Lions.
Jaxon Smolik's 47-yard deep shot to Lyrick Samuel in the second quarter proved the difference in this year's spring game.
Lambert-Smith had a bounce-back campaign after transferring to Auburn.
The Nittany Lions punched their ticket to Ann Arbor for the Big Ten Tournament semifinals.
Pena made second team All-ACC in 2024.
Ward became the first Nittany Lion to find a new home this spring.