Knowles was announced as Penn State's next defensive coordinator on January 27.
What the two Nittany Lions now have in common: Penn State degrees, Super Bowl rings, and Matty Fresh songs.
The Joe Humphreys Fly Fishing program has a storied history at Penn State.
The Nittany Lions have dropped seven of their last eight games and are in danger of missing the Big Ten Tournament.
Penn State Athletics reported $5.6 million in profit for the 2024 fiscal year.
Knowles comes to Happy Valley with 36 years of coaching experience.