Wilson started in 22 games in three seasons at Harvard and was named to the All-Ivy League second team in 2019
Three Nittany Lions are officially headed to Super Bowl LV.
Jones appeared in five games for the Nittany Lions this season.
O'Brien was an early candidate for the job after Steve Sarkisian left Tuscaloosa for the head coaching spot at Texas.
Barnwell was the only player committed to Penn State's 2023 class.
Bowens is the third Nittany Lion to transfer away from the program this offseason, joining Antonio Shelton and Judge Culpepper.