Prior to Saturday, Penn State’s last football loss came 323 days ago, when they fell to Indiana 27-24. But after blowing a 17-0 lead to Wisconsin, that streak has ended.

The 24-20 upset defeat knocked the Nittany Lions out of the AP Top 25 and drastically decreased their odds of making the College Football Playoff. The offense was catastrophically bad, scoring just one touchdown on 13 total drives.

Here’s a quick review of each position group’s performance against the Badgers.

Quarterbacks: F

Rocco Becht struggled mightily against Wisconsin, completing just 12 of his 29 pass attempts for 129 yards and an interception.

He also tacked on 10 rushes for 23 yards, including a three-yard dash for a touchdown in the second quarter.

Running Backs: D-

Penn State’s ground game continued its roller coaster against Wisconsin. Four runners combined for just 96 yards on 26 carries.

James Peoples had the only respectable day, rushing 14 times for 67 yards and averaging 4.8 yards per carry. Quinton Martin Jr. followed with six carries for 14 yards, then Carson Hansen with four carries for 11 yards. Cam Wallace, who rushed for 77 yards against Buffalo last week, totted the ball just twice for 4 yards.

Even while up 17-0 and 20-10, the Nittany Lions couldn’t lean on the rushing attack to milk the clock, sustain drives, and get out of Week 4 unscathed.

Receivers: F

The locker room expressed confidence before the game that Koby Howard’s absence wouldn’t be world-ending for Penn State’s offense. Unfortunately, that proved untrue, as Chase Sowell was the only receiver to log a catch against the Badgers. He caught six of his nine targets for 70 yards.

Notably, Brett Eskildsen could not reel in any of his six targets while Koby Howard was out. Zay Robinson, who was expected to see an increased workload due to Howard’s injury, saw just one target that he couldn’t bring in.

Penn State spent the first three weeks showcasing a receiver room that finally seemed equipped to beat coverage downfield and spark the offense, unlike the last handful of years.

However, they failed their first test against true competition, albeit without the alpha of the room.

Tight Ends: D-

Similarly, Benjamin Brahmer was the only tight end to record a reception, logging four grabs for 36 yards on eight targets. Gabe Burkle was the only other tight end to see a target.

Andrew Rappleyea was banged up in the Buffalo game and listed as a game-time decision on Saturday’s availability report.

Offensive Line: C

Brock Riker was also a game-time decision but started at center for the blue and white, helping the unit surrender just two quarterback hurries to the Badgers.

However, it struggled to open holes on the ground, as reflected in the running backs’ poor production. They were better in short-yardage situations, converting four of five opportunities for a first down or a touchdown.

Front Seven: B+

Penn State’s linebackers and defensive line did an exceptional job stopping the elusive Colton Joseph from making plays with his legs. Joseph dashed 13 times for 39 yards, but did punch in a score right before halftime.

Wisconsin’s runners certainly capitalized on the heightened focus on Joseph, as Abu Sama III and Darrion Dupree each averaged over 6.5 yards per carry.

Sama totted it five times for 34 yards and Dupree eight times for 52 yards. Neither scored, but both made chain-moving plays that cost the Nittany Lions when it mattered.

Linebacker Tony Rojas bolstered the group’s body of work by continuing his takeaway trend with an interception in the fourth quarter. The redshirt junior also had a sack and two tackles for loss.

Overall, there wasn’t much more the front seven could have done on Saturday night.

Secondary: A-

Penn State’s defensive backs picked Joseph off twice, thanks to Josiah Zayas and Daryus Dixson.

They allowed 235 passing yards but only allowed Joseph to complete 18 of his 38 passes. It’s even more impressive when you consider a blatant offensive pass interference that let Jacob Harris get free for a 72-yard go-ahead touchdown. The passing numbers allowed should have been even smaller.

The unit allowed a touchdown to Eugene Hilton Jr. with just over three minutes to play, but outside of that, they executed their assignments.

Special Teams: A

Ryan Barker was perfect, guiding through two field goals and two extra points.

The punting unit was busy on Saturday, as Nathan Tiyce booted seven points for 299 yards. He averaged 42.7 yards per punt and pinned the Badgers inside their own 20-yard line twice.

Robinson maintained his strong return presence with a 20-yard kickoff return and two punt returns for 11 and five yards.